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We study Lefschetz pencils on symplectic four-manifolds via the associated spheres in
the moduli spaces of curves, and in particular their intersections with certain natural
divisors. An invariant defined from such intersection numbers can distinguish
manifolds with torsion first Chern class. We prove that pencils of large degree always
give spheres which behave ‘homologically’ like rational curves; contrastingly, we give
the first constructive example of a symplectic non-holomorphic Lefschetz
pencil. We also prove that only finitely many values of signature or Euler
characteristic are realised by manifolds admitting Lefschetz pencils of genus two
curves.
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